It's actually way more complicated than the article claims, it mistakes correlation for causality, and is self justifying with a pseudoscientific claim.
Pseudoscientific claim : sugar content of fruits has increased due to selective breeding GENERALLY FALSE:
The majority of fruits (aside from apple, apricot and peach) have not seen their sugar content increase in cultivation, but rather their size; in fact most modern cultivars have a lower sugar content (and flavour!) than their wild counterpart.
Which leads to
mistaking correlation for causality :
animals becomes fat because of fruits YES AND NO :
It's rather the fact that zoo animals eat too much sugar in comparison with their activity level, in other words, zoo animals are bored, so they eat and do not spend energy, hence they get fat, exactly like a kid sitting in front of TV all day with food, it's not the fruits the problem, it's the way of life, but it's easier to claim that the fruits are the cause of the issue rather than admitting that it's a direct consequence of the conditions of life in zoos (which have to be improved).