It takes about a month or so for mature males hormones to decrease AFTER the neuter, it's not an instant kind of thing.
On one of your previous posts May 5th was right after the neuter, it hasn't been quit a month yet. So if you've been trying to intro between then and now that is one reason they aren't gelling.
The hormone level in Desmond is still to high, as his scent gland on top of his head shrinks, his hormones decrease which should be leveling off in the next couple of weeks.
Cages shouldn't be any closer than 12 inches to prevent tail injuries. Their tails are about 6 or 7 inches long, their arm reach from inside the cage is about 3 inches if a tail in one cage is sticking out a glider in another can grab it.
* depending on the cage type.
You have another option, you have 2 girls and 2 neutered boys. You could intro one of the girls to Desmond. He was with them once and they are female.....so that intro should go easier than the 2 boys.
Then you'd still have 2 cages and 2 play times but no single glider. You may have gotten Harrison specifically for Desmond, but gliders have a way of throwing a wrench in the best made plans. If you go this route, let the new pairs be for awhile with continued scent and pouch swapping between the 2 pairs.
Then in a month or two you could try to intro the 2 pairs to have one colony of 4. By then Desmond's hormone will have leveled off and that won't be working against you.
If the boys still don't get a long at that point you may have to come to the understanding that they just aren't going to. But I think that will change once the hormones level off.
Or You can UP your scent swapping game. Let the sleep pouches get good and dirty before you swap them, like smellyyyyyy dirty don't wash them for a couple weeks. Then swap them but don't just put the pouch in the cage. Put the gliders from cage 1 in the pouch from cage 2 (and vice versa) and rub the pouch alll over them (Sort of like washing dedicates by hand) you WANT the stink from the pouch rubbed onto the gliders in both cages. Do this Every time you swap the pouch to the opposite cage.
Especially do this right before intro's it helps. This makes 2 cages with their own distinct smell, one new smell of them combined so the gliders themselves all have the same scent.